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After installing second Exch2007 server, mailboxes cannot send outside of domain

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After installing second Exch2007 server, mailboxes cann... - 15.Nov.2016 11:59:22 AM

Rugged311

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Joined: 15.Nov.2016
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As part of a migration to Exchange 2013, I've added a second Exchange 2007 server to my domain. After installation-- during which I installed Mailbox, Client Access, and Hub transport roles--I moved some test mailboxes to the new server. The test mailboxes can send/receive from other mailboxes on the old server, and can RECEIVE from outside the domain, but cannot SEND emails to outside addresses.

After ~24 hours I receive a delivery delay, and finally an Undeliverable error from the second server with a 4.4.7 QUEUE.Expired message.

My understanding was that the two Exchange servers would detect the AD set-up and work together, so I didn't make any changes to any send/receive connectors, and they appear identical. I also haven't done anything with external DNS or NAT, as I understand everything flows through the original server.

First Exch2007 server is SP1 RU10 (I can't remove this and install SP3, which is why I decided to build a second Exchange server). Server OS is Server 2003 R2 SP2

Second Exch2007 server is SP3. Server OS is Server 2008 R2 Sp1